Mr Mneary,
The current for the engine passes from the the DCC booster, across the track through the engine, through a shorted bridge rectifier (two heavy duty diodes in series for each half DCC cycle) and back to the dcc Booster.
The diodes, when conducting, provide a maximum voltage of 0.7 volts each (1.4 volts total) across the two back to back base/emitter junctions of the Twin-T transistors. i.e Most of the engine+car current passes through the two power diodes, allowing signal transistors to be used for the detectors (Twin-T)